Orion Pax-the former Optimus Prime-follows a rogue group of Decepticons into one of the greatest mysteries ever to face the Autobots! Also, the mysterious Metrotitan arrives on Cybertron-smashing into the ruins of the legendary Crystal City! Meanwhile, violence breaks out in the heart of the city...and Prowl looks for someone to blame. Is Dirge a convenient target-or the key to what comes next?

Collects issues #10–11 and the 2012 Annual.